Mictlantecuhtli, also known as the Presser of Destruction is a bat-like demon that lives in the deepest caves of inferno and serves as a summon in Bayonetta 2. It is said to carry plague wherever it goes, it is also able to access an understanding of the world that no creature relying on its sight could see.
Book of Infernal Demons
"A gigantic bat that lurks in one of the darkest and deepest of Inferno's endless caves. It is said to leave trails of the plague wherever it goes.
The protrusion on its head is not an eye, but a spiritual sensory organ that can allow Mictlantecuhtli to access the world in far deeper ways than beings who must only rely on sight alone."